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Threshold is a suspenseful drama about a team of experts who are assembled when the US Navy makes a chillling discovery: an extraterrestrial craft has landed in the mid-Atlantic ocean. Dr. Molly Caffrey is a contingency analyst whose job it is to devise response plans for worst-case scenarios including threats to national security. In a single instant her life changes when one of those plans, Threshold, is activated by Deputy National Security Advisor J. T. Baylock.

Armed only with her hypothetical strategy to address the appearance of the aliens on Earth, Caffrey now finds herself thrust into the midst of a global crisis. She hand-picks a team of specialists to prepare for first contact: Dr. Nigel Fenway, a disillusioned NASA microbiologist; Lucas Pegg, a brilliant but neurotic physicist; and Cavennaugh, a highly trained covert operative with a mysterious past. Together they decipher the intent of the craft, the fate of the ship's crew and begin preparations for the worst case scenario - an alien invasion.